{"id":238,"date":"2017-01-23T16:36:00","date_gmt":"2017-01-23T16:36:00","guid":{"rendered":"https:\/\/pressbooks.ccconline.org\/introduction-to-oceanography\/chapter\/7-1-primary-production\/"},"modified":"2021-10-25T18:20:14","modified_gmt":"2021-10-25T18:20:14","slug":"7-1-primary-production","status":"publish","type":"chapter","link":"https:\/\/pressbooks.ccconline.org\/introduction-to-oceanography\/chapter\/7-1-primary-production\/","title":{"raw":"7.1 Primary Production","rendered":"7.1 Primary Production"},"content":{"raw":"[pb_glossary id=\"1603\"]Primary production[\/pb_glossary] is the creation of new organic matter from inorganic substrates, and it is this organic matter that serves as the base of the food web for most marine consumers. Primary production generally refers to the process of <strong>[pb_glossary id=\"1024\"]photosynthesis[\/pb_glossary],<\/strong> or the utilization of light energy to produce chemical fuels that is undertaken by plants and algae according to the reaction:\n<p style=\"text-align: center\">6CO<sub>2<\/sub> + 6H<sub>2<\/sub>O \u2192 C<sub>6<\/sub>H<sub>12<\/sub>O<sub>6<\/sub> + 6O<sub>2<\/sub><\/p>\nHere, powered by light energy, carbon dioxide and water combine to produce glucose and oxygen. However, primary production is also carried out by bacteria in the absence of light through <strong>[pb_glossary id=\"618\"]chemosynthesis[\/pb_glossary]<\/strong>. Instead of light providing the energy for the reaction, the energy comes from the oxidation of inorganic materials, such as hydrogen sulfide (see <a href=\"\/chapter\/4-11-hydrothermal-vents\/\">section 4.11<\/a> on [pb_glossary id=\"846\"]hydrothermal vents[\/pb_glossary]). Here we will concentrate on photosynthesis because it plays a much larger role in total oceanic productivity than chemosynthesis.\n\nThe organisms responsible for oceanic primary production include a wide diversity of marine plants and algae. While many people may be more familiar with the larger seagrasses and macroalgae (seaweeds), by far the greatest amount of photosynthesis in the ocean comes from microscopic algae, the <strong>[pb_glossary id=\"1028\"]phytoplankton[\/pb_glossary]<\/strong>. The term \"[pb_glossary id=\"1036\"]plankton[\/pb_glossary]\" refers to organisms that drift with the currents, and the phytoplankton are the free-floating algae that undergo photosynthesis (contrast this with the <strong>[pb_glossary id=\"1094\"]zooplankton[\/pb_glossary]<\/strong>, who are the drifting animals). In <a href=\"\/chapter\/7-2-the-producers\/\">section 7.2<\/a> we will take a closer look at the organisms responsible for oceanic primary production.\n\nThe total amount of organic material created by the producers is called the <strong>[pb_glossary id=\"796\"]gross primary productivity[\/pb_glossary]<\/strong>, or <strong>total production<\/strong>. However, the primary producers consume a portion of this organic matter themselves through respiration, so the total amount that is left to support the consumers in the ocean is called <strong>[pb_glossary id=\"968\"]net production[\/pb_glossary]<\/strong> (gross productivity - respiration). Gross production can be divided into two components, new production and regenerated production. <strong>[pb_glossary id=\"970\"]New production[\/pb_glossary]<\/strong> is supported by [pb_glossary id=\"980\"]nutrients [\/pb_glossary] brought in from outside of the local ecosystem through processes such as [pb_glossary id=\"1244\"]upwelling [\/pb_glossary] or ocean currents. <strong>[pb_glossary id=\"1074\"]Regenerated production[\/pb_glossary]<\/strong> results from the recycling of nutrients within an ecosystem.\n\nOverall, marine productivity is similar to terrestrial production. Marine net production is about 35-50 billion metric tons per year, while terrestrial production reaches 50-70 billion tons per year. However, the biomass responsible for that production in the ocean is about 1-2 billion metric tons, compared to 600-1000 billion metric tons of biomass in terrestrial systems. So the oceans are producing almost as much organic material as terrestrial producers, but are doing it from only a fraction of the amount of producer biomass. One reason for this discrepancy is that the phytoplankton are constantly being consumed, while much of the terrestrial biomass is much longer-lived than the plankton.","rendered":"<p>Primary production is the creation of new organic matter from inorganic substrates, and it is this organic matter that serves as the base of the food web for most marine consumers. Primary production generally refers to the process of <strong><a class=\"glossary-term\" aria-haspopup=\"dialog\" aria-describedby=\"definition\" href=\"#term_238_1024\">photosynthesis<\/a>,<\/strong> or the utilization of light energy to produce chemical fuels that is undertaken by plants and algae according to the reaction:<\/p>\n<p style=\"text-align: center\">6CO<sub>2<\/sub> + 6H<sub>2<\/sub>O \u2192 C<sub>6<\/sub>H<sub>12<\/sub>O<sub>6<\/sub> + 6O<sub>2<\/sub><\/p>\n<p>Here, powered by light energy, carbon dioxide and water combine to produce glucose and oxygen. However, primary production is also carried out by bacteria in the absence of light through <strong><a class=\"glossary-term\" aria-haspopup=\"dialog\" aria-describedby=\"definition\" href=\"#term_238_618\">chemosynthesis<\/a><\/strong>. Instead of light providing the energy for the reaction, the energy comes from the oxidation of inorganic materials, such as hydrogen sulfide (see <a href=\"\/chapter\/4-11-hydrothermal-vents\/\">section 4.11<\/a> on <a class=\"glossary-term\" aria-haspopup=\"dialog\" aria-describedby=\"definition\" href=\"#term_238_846\">hydrothermal vents<\/a>). Here we will concentrate on photosynthesis because it plays a much larger role in total oceanic productivity than chemosynthesis.<\/p>\n<p>The organisms responsible for oceanic primary production include a wide diversity of marine plants and algae. While many people may be more familiar with the larger seagrasses and macroalgae (seaweeds), by far the greatest amount of photosynthesis in the ocean comes from microscopic algae, the <strong><a class=\"glossary-term\" aria-haspopup=\"dialog\" aria-describedby=\"definition\" href=\"#term_238_1028\">phytoplankton<\/a><\/strong>. The term &#8220;<a class=\"glossary-term\" aria-haspopup=\"dialog\" aria-describedby=\"definition\" href=\"#term_238_1036\">plankton<\/a>&#8221; refers to organisms that drift with the currents, and the phytoplankton are the free-floating algae that undergo photosynthesis (contrast this with the <strong><a class=\"glossary-term\" aria-haspopup=\"dialog\" aria-describedby=\"definition\" href=\"#term_238_1094\">zooplankton<\/a><\/strong>, who are the drifting animals). In <a href=\"\/chapter\/7-2-the-producers\/\">section 7.2<\/a> we will take a closer look at the organisms responsible for oceanic primary production.<\/p>\n<p>The total amount of organic material created by the producers is called the <strong><a class=\"glossary-term\" aria-haspopup=\"dialog\" aria-describedby=\"definition\" href=\"#term_238_796\">gross primary productivity<\/a><\/strong>, or <strong>total production<\/strong>. However, the primary producers consume a portion of this organic matter themselves through respiration, so the total amount that is left to support the consumers in the ocean is called <strong><a class=\"glossary-term\" aria-haspopup=\"dialog\" aria-describedby=\"definition\" href=\"#term_238_968\">net production<\/a><\/strong> (gross productivity &#8211; respiration). Gross production can be divided into two components, new production and regenerated production. <strong><a class=\"glossary-term\" aria-haspopup=\"dialog\" aria-describedby=\"definition\" href=\"#term_238_970\">New production<\/a><\/strong> is supported by <a class=\"glossary-term\" aria-haspopup=\"dialog\" aria-describedby=\"definition\" href=\"#term_238_980\">nutrients <\/a> brought in from outside of the local ecosystem through processes such as <a class=\"glossary-term\" aria-haspopup=\"dialog\" aria-describedby=\"definition\" href=\"#term_238_1244\">upwelling <\/a> or ocean currents. <strong><a class=\"glossary-term\" aria-haspopup=\"dialog\" aria-describedby=\"definition\" href=\"#term_238_1074\">Regenerated production<\/a><\/strong> results from the recycling of nutrients within an ecosystem.<\/p>\n<p>Overall, marine productivity is similar to terrestrial production. Marine net production is about 35-50 billion metric tons per year, while terrestrial production reaches 50-70 billion tons per year. However, the biomass responsible for that production in the ocean is about 1-2 billion metric tons, compared to 600-1000 billion metric tons of biomass in terrestrial systems. So the oceans are producing almost as much organic material as terrestrial producers, but are doing it from only a fraction of the amount of producer biomass. One reason for this discrepancy is that the phytoplankton are constantly being consumed, while much of the terrestrial biomass is much longer-lived than the plankton.<\/p>\n<div class=\"glossary\"><span class=\"screen-reader-text\" id=\"definition\">definition<\/span><template id=\"term_238_1603\"><div class=\"glossary__definition\" role=\"dialog\" data-id=\"term_238_1603\"><div tabindex=\"-1\"><\/div><button><span aria-hidden=\"true\">&times;<\/span><span class=\"screen-reader-text\">Close definition<\/span><\/button><\/div><\/template><template id=\"term_238_1024\"><div class=\"glossary__definition\" role=\"dialog\" data-id=\"term_238_1024\"><div tabindex=\"-1\"><p>the production of organic compounds from carbon dioxide and water, using sunlight as an energy source (5.5)<\/p>\n<\/div><button><span aria-hidden=\"true\">&times;<\/span><span class=\"screen-reader-text\">Close definition<\/span><\/button><\/div><\/template><template id=\"term_238_618\"><div class=\"glossary__definition\" role=\"dialog\" data-id=\"term_238_618\"><div tabindex=\"-1\"><p>the creation of organic compounds using the energy from inorganic chemical reactions (4.11)<\/p>\n<\/div><button><span aria-hidden=\"true\">&times;<\/span><span class=\"screen-reader-text\">Close definition<\/span><\/button><\/div><\/template><template id=\"term_238_846\"><div class=\"glossary__definition\" role=\"dialog\" data-id=\"term_238_846\"><div tabindex=\"-1\"><p>area of the seafloor where superheated water seeps out of the crust (4.11)<\/p>\n<\/div><button><span aria-hidden=\"true\">&times;<\/span><span class=\"screen-reader-text\">Close definition<\/span><\/button><\/div><\/template><template id=\"term_238_1028\"><div class=\"glossary__definition\" role=\"dialog\" data-id=\"term_238_1028\"><div tabindex=\"-1\"><p>drifting, usually single-celled algae that undergo photosynthesis (7.1)<\/p>\n<\/div><button><span aria-hidden=\"true\">&times;<\/span><span class=\"screen-reader-text\">Close definition<\/span><\/button><\/div><\/template><template id=\"term_238_1036\"><div class=\"glossary__definition\" role=\"dialog\" data-id=\"term_238_1036\"><div tabindex=\"-1\"><p>an organism that cannot swim effectively, so it drifts with the currents (7.1)<\/p>\n<\/div><button><span aria-hidden=\"true\">&times;<\/span><span class=\"screen-reader-text\">Close definition<\/span><\/button><\/div><\/template><template id=\"term_238_1094\"><div class=\"glossary__definition\" role=\"dialog\" data-id=\"term_238_1094\"><div tabindex=\"-1\"><p>small, drifting carnivorous organisms (7.1)<\/p>\n<\/div><button><span aria-hidden=\"true\">&times;<\/span><span class=\"screen-reader-text\">Close definition<\/span><\/button><\/div><\/template><template id=\"term_238_796\"><div class=\"glossary__definition\" role=\"dialog\" data-id=\"term_238_796\"><div tabindex=\"-1\"><p>the total amount of organic material created by primary producers (7.1)\u00a0<strong><br \/>\n<\/strong><\/p>\n<\/div><button><span aria-hidden=\"true\">&times;<\/span><span class=\"screen-reader-text\">Close definition<\/span><\/button><\/div><\/template><template id=\"term_238_968\"><div class=\"glossary__definition\" role=\"dialog\" data-id=\"term_238_968\"><div tabindex=\"-1\"><p>total primary production minus the organic compounds used up by respiration by the producers (7.1)<\/p>\n<\/div><button><span aria-hidden=\"true\">&times;<\/span><span class=\"screen-reader-text\">Close definition<\/span><\/button><\/div><\/template><template id=\"term_238_970\"><div class=\"glossary__definition\" role=\"dialog\" data-id=\"term_238_970\"><div tabindex=\"-1\"><p>primary production supported by nutrients brought in from outside of the local ecosystem (7.1)<\/p>\n<\/div><button><span aria-hidden=\"true\">&times;<\/span><span class=\"screen-reader-text\">Close definition<\/span><\/button><\/div><\/template><template id=\"term_238_980\"><div class=\"glossary__definition\" role=\"dialog\" data-id=\"term_238_980\"><div tabindex=\"-1\"><p>in the context of primary production, substances required by photosynthetic organisms to undergo growth and reproduction (5.6)<\/p>\n<\/div><button><span aria-hidden=\"true\">&times;<\/span><span class=\"screen-reader-text\">Close definition<\/span><\/button><\/div><\/template><template id=\"term_238_1244\"><div class=\"glossary__definition\" role=\"dialog\" data-id=\"term_238_1244\"><div tabindex=\"-1\"><p>process by which deeper water is brought to the surface (9.5)<\/p>\n<\/div><button><span aria-hidden=\"true\">&times;<\/span><span class=\"screen-reader-text\">Close definition<\/span><\/button><\/div><\/template><template id=\"term_238_1074\"><div class=\"glossary__definition\" role=\"dialog\" data-id=\"term_238_1074\"><div tabindex=\"-1\"><p>primary production resulting from the recycling of nutrients within an ecosystem (7.1)<\/p>\n<\/div><button><span aria-hidden=\"true\">&times;<\/span><span class=\"screen-reader-text\">Close definition<\/span><\/button><\/div><\/template><\/div>","protected":false},"author":33,"menu_order":41,"template":"","meta":{"pb_show_title":"on","pb_short_title":"","pb_subtitle":"","pb_authors":["paul-webb"],"pb_section_license":"cc-by"},"chapter-type":[],"contributor":[60],"license":[52],"class_list":["post-238","chapter","type-chapter","status-publish","hentry","contributor-paul-webb","license-cc-by"],"part":234,"_links":{"self":[{"href":"https:\/\/pressbooks.ccconline.org\/introduction-to-oceanography\/wp-json\/pressbooks\/v2\/chapters\/238","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/pressbooks.ccconline.org\/introduction-to-oceanography\/wp-json\/pressbooks\/v2\/chapters"}],"about":[{"href":"https:\/\/pressbooks.ccconline.org\/introduction-to-oceanography\/wp-json\/wp\/v2\/types\/chapter"}],"author":[{"embeddable":true,"href":"https:\/\/pressbooks.ccconline.org\/introduction-to-oceanography\/wp-json\/wp\/v2\/users\/33"}],"version-history":[{"count":2,"href":"https:\/\/pressbooks.ccconline.org\/introduction-to-oceanography\/wp-json\/pressbooks\/v2\/chapters\/238\/revisions"}],"predecessor-version":[{"id":1323,"href":"https:\/\/pressbooks.ccconline.org\/introduction-to-oceanography\/wp-json\/pressbooks\/v2\/chapters\/238\/revisions\/1323"}],"part":[{"href":"https:\/\/pressbooks.ccconline.org\/introduction-to-oceanography\/wp-json\/pressbooks\/v2\/parts\/234"}],"metadata":[{"href":"https:\/\/pressbooks.ccconline.org\/introduction-to-oceanography\/wp-json\/pressbooks\/v2\/chapters\/238\/metadata\/"}],"wp:attachment":[{"href":"https:\/\/pressbooks.ccconline.org\/introduction-to-oceanography\/wp-json\/wp\/v2\/media?parent=238"}],"wp:term":[{"taxonomy":"chapter-type","embeddable":true,"href":"https:\/\/pressbooks.ccconline.org\/introduction-to-oceanography\/wp-json\/pressbooks\/v2\/chapter-type?post=238"},{"taxonomy":"contributor","embeddable":true,"href":"https:\/\/pressbooks.ccconline.org\/introduction-to-oceanography\/wp-json\/wp\/v2\/contributor?post=238"},{"taxonomy":"license","embeddable":true,"href":"https:\/\/pressbooks.ccconline.org\/introduction-to-oceanography\/wp-json\/wp\/v2\/license?post=238"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}